Disputed term/author/ism | Author |
Entry |
Reference |
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Evidence | Darwin | Gould I 189 Evidence/Darwin/Gould: Darwin represented the view that the fossil finds were extremely incomplete. The history of most fossil species includes two characteristics that are particularly difficult to reconcile with the theory of gradual change. I 191 With the "sympatric" speciation, new forms appear within the distribution area of the previous form. The situation is different on the periphery: isolated small populations here are much more exposed to selection pressure because the periphery marks the limit of the ecological tolerance of the previous living beings. Question: What should fossil finds contain if the majority of evolution takes place in small marginalized groups? I 210 Evidence/Method/Gould: Events which, according to traditional wisdom, cannot "occur" are rarely recognized on the basis of collected evidence, they require a mechanism for their explanation after they can occur. The empirical evidence of that time seems plausible to us today, at that time there were not, because no acceptable force could be named, which would have been able to move ((s) e.g.) whole continents. >Method, >Evolution, >Justification, >Confirmation. Gould IV 33 Evidence/Darwin/Gould: Darwin wrote "The Descent of Man" in 1871(1), at that time there were no fossils, except for the Neanderthal man who is a race of our own kind, neither an ancestor nor a missing link. Gould IV 35 Explanation/evidence/Science/Darwin/Gould: how do we know that evolution controls the world and not any other principle? Darwin's answer was paradoxical. He was not looking for perfection, but for inconsistencies! There are structures and behaviours that endanger the meaningful form of an organism, but ultimately ensure its success. They increase productive efficiency. For example, the tail feathers of the peacocks and the antlers of the moose. They even weaken the species, but still secure the transmission of the genes. This can only be explained by Darwin: in addition to adaptation: sexual selection. IV 36 Sexual selection: competition of the males among each other or election by the females. Thus, the struggle for existence of the individuals drives evolution! It must be exceptionally effective, it is often capable of overcoming other equipment benefits and developments. 1. Ch. Darwin, The descent of man, and selection in relation to sex. John Murray, London 1871. |
Gould I Stephen Jay Gould The Panda’s Thumb. More Reflections in Natural History, New York 1980 German Edition: Der Daumen des Panda Frankfurt 2009 Gould II Stephen Jay Gould Hen’s Teeth and Horse’s Toes. Further Reflections in Natural History, New York 1983 German Edition: Wie das Zebra zu seinen Streifen kommt Frankfurt 1991 Gould III Stephen Jay Gould Full House. The Spread of Excellence from Plato to Darwin, New York 1996 German Edition: Illusion Fortschritt Frankfurt 2004 Gould IV Stephen Jay Gould The Flamingo’s Smile. Reflections in Natural History, New York 1985 German Edition: Das Lächeln des Flamingos Basel 1989 |
Evolution | Darwin | Gould II 329 Darwin: 2 stages of evolution 1) Random range of variation (production of raw material) 2) Selection as a conventional force. Gould II 331 The official definition of evolution/Gould: Def Evolution: "Change of gene frequencies in populations". (The process of random increase or decrease in the gene frequency is called Def "genetic drift".) The new theory of neutralism suggests that many, if not most, of the genes in individual populations owe their frequency primarily to chance. --- Mayr I 235 Darwin (early): change by adaption. - Vs: adaption can never explain the enormous diversity of organic life, because there is no increase in the number of species. Mayr I 236 Darwin/Mayr: The Origin of Species: 5 Main Theories 1) Organisms are constantly evolving over time (evolution as such). 2) Different types of organisms are derived from a common ancestor. 3) Species multiply over time (speciation) 4) Evolution takes place in the form of gradual change. (GradualismVsSaltationism). 5) The evolutionary mechanism consists in the competition of subordinate unique individuals for limited resources that lead to differences in survival and reproduction (natural selection). >Selection. --- Gould IV 357 Evolution/Darwin: Thesis: no inner dynamics drives life forward! If the environment really did not change, it would not be impossible for evolution to come to a standstill crunchingly! Actually, the species lead their lives independently of each other! Their most important battles take place against climate, geological and geographical conditions. (>Species). |
Gould I Stephen Jay Gould The Panda’s Thumb. More Reflections in Natural History, New York 1980 German Edition: Der Daumen des Panda Frankfurt 2009 Gould II Stephen Jay Gould Hen’s Teeth and Horse’s Toes. Further Reflections in Natural History, New York 1983 German Edition: Wie das Zebra zu seinen Streifen kommt Frankfurt 1991 Gould III Stephen Jay Gould Full House. The Spread of Excellence from Plato to Darwin, New York 1996 German Edition: Illusion Fortschritt Frankfurt 2004 Gould IV Stephen Jay Gould The Flamingo’s Smile. Reflections in Natural History, New York 1985 German Edition: Das Lächeln des Flamingos Basel 1989 Mayr I Ernst Mayr This is Biology, Cambridge/MA 1997 German Edition: Das ist Biologie Heidelberg 1998 |
Evolution | Gould | Dennett I 412 Evolution/Gould theory: the key difference in evolution is not simple adaptation but speciation. (DennettVs). Gould: thesis: species are fragile but have unalterable structures. There are no improvements in species, only closed discarding. Correct level: the correct level are not the genes but entire species or clades. Species/Gould/(s): species are not going to be improved, but discarded. Level/explanation/Dennett: as software/hardware: some is better explained on one level, others is better explained on a different level. >Explanation, >Darwinism. Gould I 88ff Evolution/Darwinism/individual/Gould: individuals do not develop evolutionary, they can only grow, reproduce and die. Evolutionary changes occur in groups of interacting organisms. Species are the units of evolution. Orthodox Darwinism/Gould: thesis: gene mutate, individuals are subject to selection and species evolve evolutionary. I 131 Evolution/Gould: Thesis: I do not imagine evolution as a ladder, but rather in the form of a shrub with many branches. Therefore: the more species the better. I 133 The importance of this point can be seen in the development of molecules. The number of differences between amino acids clearly correlates with the time since the diversion of development lines. The longer the separation, the greater the differences. This is how a molecular clock was developed. The Darwinians were generally surprised by the regularity of this clock. After all, the selection should proceed at a noticeably different speed for the different development lines at different times. I 134 VsDarwinism: the Darwinists are actually forced to contemplate that the regular molecular clock represents an evolution that is not subject to selection, but to the random fixation of neutral mutations. We have never been able to separate ourselves from the concept of the evolution of the human being, which puts the brain in the centre of attention. The Australopithecus afarensis disproved what had been predicted by astute evolutionary theorists such as Ernst Haeckel and Friedrich Engels. Tradition: general view: that the upright gait represented an easily attainable gradual development, and the increase in brain volume represented a surprisingly rapid leap. I 136 GouldVs: I would like to take the opposite view: in my opinion, the upright gait is a surprise, a difficult event to achieve, a rapid and fundamental transformation of our anatomy. In anatomical terms, the subsequent enlargement of our brain is a secondary epiphenomenon, a simple transition embedded in the general pattern of human evolution. Bipedality is not an easy achievement, it represents a fundamental transformation of our anatomy, especially of the feet and pelvis. I 191 Evolution/Gould: evolution essentially proceeds in two ways: a) Definition phyletic transformation: an entire population changes from one state to another. If all evolutionary changes were to occur in this way, life would not last long. This is because a phyletic transformation does not lead to an increase in diversity and variety, only to a transformation from one state to another. Now that extinction (by eradication) is so widespread, everything that does not have the ability to adapt would soon be destroyed. b) Definition speciation: new species branch off from existing ones. All speciation theories assume that splits occur quickly in very small populations. With the "sympatric" speciation, new forms appear within the distribution area of the previous form. Large stable central populations have a strong homogenizing influence. New mutations are impaired by the strong previous forms: they may slowly increase in frequency, but a changed environment usually reduces their selective value long before they can assert themselves. Thus, a phyletic transformation of the large populations should be very rare, as the fossil finds prove. It looks different in the periphery: isolated small populations here are much more exposed to the selection pressure, because the periphery marks the limit of the ecological tolerance of the previous living beings. I 266 Evolution/Biology/Gould: evolution proceeds by replacing the nucleotides. II 243 Evolution/Gould: thesis: evolution has no tendency. II 331 Evolution/Gould: official definition of evolution/Gould: evolution is the "change of gene frequencies in populations". (The process of random increase or decrease of the gene frequency is called definition "genetic drift".) The new theory of neutralism suggests that many, if not most, genes in individual populations owe their frequency primarily to chance. IV 199 Evolution/species richness: the change from a few species and many groups to a few groups and many species would occur even in the case of purely coincidental extinction if every speciation process at the beginning of life's history had been accompanied by average major changes. IV 221 Evolution/Gould: pre-evolutionary theory: a pre-evolutionary theory is "the chain of being": it is the old idea that every organism is a link. It confuses evolution with higher development and has been misinterpreted as a primitive form of evolution, but has nothing to do with it! The thesis is emphatically antievolutionary. Problem: there are no links between vertebrates and invertebrates IV 223 Intermediate form: the theory assumed asbestos as an intermediate form between minerals and plants due to the fibrous structure. Hydra and corals were seen as an intermediate form between plants and animals. (Today: both are animals of course.) Absurd: it is absurd to assume a similarity between plants and baboons, because plants lose their leaves and baboon babies lose their hair. IV 346 Evolution/Gould: evolution is not developing in the direction of complexity, why should it? |
Gould I Stephen Jay Gould The Panda’s Thumb. More Reflections in Natural History, New York 1980 German Edition: Der Daumen des Panda Frankfurt 2009 Gould II Stephen Jay Gould Hen’s Teeth and Horse’s Toes. Further Reflections in Natural History, New York 1983 German Edition: Wie das Zebra zu seinen Streifen kommt Frankfurt 1991 Gould III Stephen Jay Gould Full House. The Spread of Excellence from Plato to Darwin, New York 1996 German Edition: Illusion Fortschritt Frankfurt 2004 Gould IV Stephen Jay Gould The Flamingo’s Smile. Reflections in Natural History, New York 1985 German Edition: Das Lächeln des Flamingos Basel 1989 Dennett I D. Dennett Darwin’s Dangerous Idea, New York 1995 German Edition: Darwins gefährliches Erbe Hamburg 1997 Dennett II D. Dennett Kinds of Minds, New York 1996 German Edition: Spielarten des Geistes Gütersloh 1999 Dennett III Daniel Dennett "COG: Steps towards consciousness in robots" In Bewusstein, Thomas Metzinger Paderborn/München/Wien/Zürich 1996 Dennett IV Daniel Dennett "Animal Consciousness. What Matters and Why?", in: D. C. Dennett, Brainchildren. Essays on Designing Minds, Cambridge/MA 1998, pp. 337-350 In Der Geist der Tiere, D Perler/M. Wild Frankfurt/M. 2005 |
Evolution | Mayr | I 43 Evolution/Mayr: Unit of evolution is the population (or species) and not the gene or the individual. (MayrVsDawkins). >Species, >Genes, >R. Dawkins, >Genes/Dawkins, >Evolution/Dawkins. Def Integron/Mayr: An integron is a system created by integration of subordinate units on a higher level. Integrons evolve by natural selection. They are adapted systems at each level because they contribute to the fitness (suitability) of an individual. >Selection. I 183 Evolution/Mayr: Species is the decisive entity of evolution. I 230 Evolution/Progress/Mayr: Cohesion: an expression of the fact that the system of development has become very narrow. Evolution: proceeds very slowly in large, member-rich species, and very quickly in small peripheral isolated groups. >Speciation, >punctuated equilibrium/Eldredge/Gould. A start-up population with few individuals and therefore little hidden genetic variation can more easily assume a different genotype. Macroevolution: is most strongly determined by the geographical factor (isolation). I 234 Evolution/Mayr: the concepts: 1) Rapid evolution: (transmutationism): type jump. Even after Darwin some researchers (including his friend Huxley) could not accept the concept of natural selection and developed saltationist theories. 2) Transformational evolution (transformationism) gradual change of the ice to the organism. Ignored by Darwin. I 235 3) Variation Evolution (Darwin) I 235 Darwin (early): adaptation modification. Vs: can never explain the enormous variety of organic life, because it does not allow for an increase in the number of species. I 236 Darwin/Mayr: The Origin of Species: 5 Main Theories 1) Organisms are constantly evolving over time (evolution as such). 2) Different species of organisms are derived from a common ancestor. 3) Species multiply over time (speciation) 4) Evolution takes the form of gradual change. (GradualismVsSaltationism). >Gradualism, >Saltationism. 5) The evolutionary mechanism consists in the competition among numerous unique individuals for limited resources that leads to differences in survival and reproduction (natural selection). I 234 Evolution/Mayr: the concepts: 1) Rapid evolution: (transmutationism): type jump. Even after Darwin some researchers (including his friend Huxley) could not accept the concept of natural selection and developed saltationist theories. 2) Transformational evolution (transformationism) gradual change of the ice to the organism. Ignored by Darwin. I 235 3) Variation Evolution (Darwin) I 235 Darwin (early): adaptation modification. Vs: can never explain the enormous variety of organic life, because it does not allow for an increase in the number of species. I 236 Darwin/Mayr: The Origin of Species: 5 Main Theories 1) Organisms are constantly evolving over time (evolution as such). 2) Different species of organisms are derived from a common ancestor. 3) Species multiply over time (speciation) 4) Evolution takes the form of gradual change. (GradualismVsSaltationism). 5) The evolutionary mechanism consists in the competition among numerous unique individuals for limited resources that leads to differences in survival and reproduction (natural selection). >Selection. I 377 Evolution of life: a chemical process involving autocatalysis and a directing factor. Prebiotic selection. Cf. >St. Kauffman. I 237 Pasteur: proofed the impossibility of life in oxygen-rich atmosphere! In 1953, Stanley Miller grew amino acids, urea and other organic molecules in a glass flask by discharging electricity into a mixture of methane, ammonium, hydrogen, and water vapor. I 238 Proteins, nucleic acids: the organisms must form these larger molecules themselves. Amino acids, pyrimidines, puridine do not need to formed by the organisms themselves. I 239 Molecular biology: discovered that the genetic code is the same for bacteria, which do not have nuclei, as in protists, fungi, animals and plants. I 240 Missing link: Archaeopteryx: half bird half reptile. Not necessarily direct ancestor. Speciation: a) dichopatric: a previously connected area is divided by a new barrier: mountain range, inlets, interruption of vegetation. b) peripatrically: new start-up population emerges outside of the original distribution area. c) sympatric speciation: new species due to ecological specialization within the area of distribution. Darwin's theory of gradualism. >Gradualism. I 243 VsGardualism: cannot explain the emergence of completely new organs. Problem: How can a rudimentary wing be enlarged by natural selection before it is suitable for flying? I 244 Darwin: two possible solutions: a) Intensification of the function: E.g. eyes, e.g. the development of the anterior limbs of moles, whales, bats. b) Functional change: E.g. Antennae of daphia (water flea): additional function of the swimming paddle, which is enlarged and modified under selection pressure. E.g. Gould: Feathers probably first for temperature control before any animal could fly. Function/Biology: Functional differences are also related to behavioral patterns, e.g. feather cleaning. Competing theories on evolutionary change I 247 Salationism: Huxley later Bateson, de Vries, (Mendelists). The saltationist emergence of new species only occurs poyploidy and some other forms of chromosomal restructuring (very rare) during sexual reproduction. Teleological theories: assume that nature has a principle: Osbron's arsitogenesis, Chardin's omega principle. Should lead to perfection. >Teilhard de Chardin. Lamarck's Theories: Changes go back to use and non-use, environmental conditions. Until the 1930s! I 248 Def "soft inheritance" (acquired characteristics). Was refuted by genetics. Def "hard inheritance" (so-called "central dogma"): the information contained in the proteins (the phenotype) cannot be passed on to the nucleic acids (the genotype)! (Insight of molecular biology). I 256 Macroevolution: after saltationism, soft heredity and autogenesis, had been refuted with evolution, macroevolution had to be explained more and more as a phenomenon on the level of the population, i.e. as a phenomenon directly attributable to events and processes during microevolution. (Speciation: faster in isolation). (>Gould, Eldredge, 1971(1): "punctuated equilibrium", punctualism.) I 281 New: we know today that the cycles of herbivores elicit those of the predators and not vice versa! Coevolution: E.g. the Yucca moth destroys the plant's ovules by its larvae, but pollens the flowers. 1. N. Eldredge, S. J. Gould: Punctuated equilibria: an alternative to phyletic gradualism. In: T. Schopf (Ed), Models in Paleobiology, 82-115, San Francisco, (1972). |
Mayr I Ernst Mayr This is Biology, Cambridge/MA 1997 German Edition: Das ist Biologie Heidelberg 1998 |
Species | Darwin | Dennett I 48 Species/Darwin: actually arbitrary! Applied to a number of individuals for convenience. Dennett I, 390ff Speciation/Darwin: Competition between closely related forms Driving force for speciation. So the presence and not the absence of two groups is therefore important. (Not "empty space in the design room"). >Species. |
Dennett I D. Dennett Darwin’s Dangerous Idea, New York 1995 German Edition: Darwins gefährliches Erbe Hamburg 1997 Dennett II D. Dennett Kinds of Minds, New York 1996 German Edition: Spielarten des Geistes Gütersloh 1999 Dennett III Daniel Dennett "COG: Steps towards consciousness in robots" In Bewusstein, Thomas Metzinger Paderborn/München/Wien/Zürich 1996 Dennett IV Daniel Dennett "Animal Consciousness. What Matters and Why?", in: D. C. Dennett, Brainchildren. Essays on Designing Minds, Cambridge/MA 1998, pp. 337-350 In Der Geist der Tiere, D Perler/M. Wild Frankfurt/M. 2005 |
Species | Dawkins | Dennett I 56 Species/Dawkins/Dennett: there is nothing that wants speciation! Before the separation, there must be a period of "temporary separation". |
Da I R. Dawkins The Selfish Gene, Oxford 1976 German Edition: Das egoistische Gen, Hamburg 1996 Da II M. St. Dawkins Through Our Eyes Only? The Search for Animal Consciousness, Oxford/New York/Heidelberg 1993 German Edition: Die Entdeckung des tierischen Bewusstseins Hamburg 1993 Dennett I D. Dennett Darwin’s Dangerous Idea, New York 1995 German Edition: Darwins gefährliches Erbe Hamburg 1997 Dennett II D. Dennett Kinds of Minds, New York 1996 German Edition: Spielarten des Geistes Gütersloh 1999 Dennett III Daniel Dennett "COG: Steps towards consciousness in robots" In Bewusstein, Thomas Metzinger Paderborn/München/Wien/Zürich 1996 Dennett IV Daniel Dennett "Animal Consciousness. What Matters and Why?", in: D. C. Dennett, Brainchildren. Essays on Designing Minds, Cambridge/MA 1998, pp. 337-350 In Der Geist der Tiere, D Perler/M. Wild Frankfurt/M. 2005 |
Species | Gould | I 223 Species/Gould: an average invertebrate species lived about 5 - 10 million years unchanged. It hardly changes in time and dies out without successors. At a higher level, evolution is basically a matter of the different successes of species and not a slow transformation of lineages. II 331 Species/Gould: Definition species: species are defined as populations isolated from all others in reproduction. When brought together with other species, they will not mix. Key question for the origin of a new species: how do isolation mechanisms develop? II 332 Traditional point of view: an originally unified population is separated from continents by drifting apart, by newly formed mountain ranges, the newly isolated groups would adapt to their new environment by adaptation. After a certain time, the populations become so different that they can no longer be crossed. New view: the ultimate success of a species may depend on the evolution of adaptation, but the act of species formation itself can be a coincidental event. Taxonomists have discovered that many groups of closely related species are not very different in form, behavior, and even in genetic equipment. However, there are striking differences in the number and shape of chromosomes, and these differences produce the isolation mechanisms that they receive as a separate species. The main change occurs in a single individual. Who should it breed with? IV 198 Species: biodiversity has certainly increased over time. Today's oceans contain at least twice as many species as the oceans in the Palaeozoic. Therefore, one could expect that they not only contain more species, but also more diverse species of organisms, with fundamentally different blueprints. But this is not the case! Today, twice as many species are put in much fewer groups of higher taxa. Today's seas are dominated by fewer groups: primarily by mussels, snails, crabs, fish and sea urchins. Each group includes many more species than any tribe in the Palaeozoic ever had. This steady decrease of organic construction types with a strong increase in the number of species is probably the most prominent trend of fossil documents! IV 199 Causality/coincidence/evolution: there may be one principle that can be identified: "initial experimentation and later standardization". For example, around 1900 there were few car brands and a much wider range of construction types. Today, there are hundreds of brands and much more uniform construction. Evolution/species richness: the change from a few species and many groups to a few groups and many species would occur even in the case of purely coincidental extinction if every speciation process at the beginning of life's history had been accompanied by average major changes. IV 201 Extinction: each is inevitable forever. An extinct experiment will never be repeated. The chances are mathematically too slim. Biologists speak of the "principle of the irreversibility of evolution". Order/coincidence/Gould: coincidental processes produce a high degree of order. The fact that they result in certain patterns does not speak against their coincidence. IV 327 Species/Gould: each species is a concatenation of improbabilities. Every species, whether human, coral or squid, is the last link in a chain that stretches back to the beginning of life. If any of these species had died out or evolved in any other direction, the end results would be very different. For example, our ancestors, the fish, developed a special fin with a stable, central bony axis. Without them, they could not have developed ashore. Nevertheless, these fins did not develop in anticipation of the necessities of rural life. They developed as adaptations to a local habitat. Necessity: human brains did not develop on a direct and necessary ascending ladder, but on winding paths full of accidents. III 264 Species/Gould: in the early days of evolution, the greatest spectrum of forms was reached, and most of the early experiments were extinct. It was accidental and not by predictable causes. Today, there are only a small number of possibilities left. >Evolution, >Explanation, >Darwinism. |
Gould I Stephen Jay Gould The Panda’s Thumb. More Reflections in Natural History, New York 1980 German Edition: Der Daumen des Panda Frankfurt 2009 Gould II Stephen Jay Gould Hen’s Teeth and Horse’s Toes. Further Reflections in Natural History, New York 1983 German Edition: Wie das Zebra zu seinen Streifen kommt Frankfurt 1991 Gould III Stephen Jay Gould Full House. The Spread of Excellence from Plato to Darwin, New York 1996 German Edition: Illusion Fortschritt Frankfurt 2004 Gould IV Stephen Jay Gould The Flamingo’s Smile. Reflections in Natural History, New York 1985 German Edition: Das Lächeln des Flamingos Basel 1989 |
Species | Lorenz | Lorenz I 110 Definition "apparant speciation"/Erik Erikson/Lorenz: Formation of differences in group-specific behavior. >Psychological theories on behavior, >Natural kind/Philosophy. |
Lorenz I K. Lorenz Das sogenannte Böse Wien 1963 |
Species | Mayr | Gould I 216 Species/Darwin/Lamarck: Species are no natural units but "purely artificial combinations"... conceptual definitions. >Order, >Systems, >Definitions, >Definability, >Classification. I 217 Species/Ernst MayrVsDarwin/MayrVsLamarck: "Species are a product of evolution and not of the human mind." >Evolution, >Evolution/Mayr. I 179 Definition Species/Mayr: device for protecting balanced, harmonic genotypes. "Biological concept of species" seeks biological reasons for the existence of species. Maybe there are other properties by chance. Biological species concept: 1. Problem: Asexual organisms do not form populations. 2. Problem: Spatial expansion with subspecies. They can become independent species in isolation over time (by acquiring new isolation mechanisms). (polytypical species). I 181 Nominalist concept of species: in nature exclusively individuals, species artificially created by humans (MayrVs: that would be arbitrariness, and nature shows that there is no arbitrariness.). I 182 Evolutionary species concept: temporal dimension, generational series of populations. MayrVs: the concept does not take into account that there are two possible ways of species development: a) Gradual transformation of a stem line into another species without altering the number of species; and b) The reproduction of species through geographical isolation. >Systems, >Definitions, >Definability. I 183 Species/Mayr: is applied to three very different objects or phenomena: 1. The species concept 2. The category species 3. The species taxa Some authors could not differentiate between them, leading to hopeless confusion in literature. Species concept: biological meaning or definition of the word "species". Category Species: certain rank in the Linnéian system. (Other categories: Order, Kingdom, Genus...) Definition Species Taxa: special populations or population groups corresponding to the species definition. They are entities ("individuals") and cannot be defined as such. Individuals cannot be defined, but can only be described and delimited. I 183 Evolution/Mayr: Species is the decisive entity of evolution. Species: a species, regardless of the individuals belonging to it, interacts as a unit with other species in the common environment. I 185 Macrotaxonomy: the classification of species (in higher-level groups) Group: mostly easily recognizable: birds, butterflies, beetles. Downward classification (actual identification). Division (aristotelian), heyday of medicinal botany. E. g. warm-blooded or non-warm-blooded - having or not having feathers. I 192 Organism types: most new types of organisms do not originate from the gradual transformation of a stem line, i. e. an existing type. Rather, a founder species penetrates into a new adaptive zone and is successful there thanks to rapid adaptive changes. For example, the more than 5000 species of songbirds are no more than the variation of a single theme. I 192 Species: the two evolutionary ways to produce a new species: a) gradual change of the phenotype and b) increasing diversity (speciation) are only loosely related. I 192 Selection pressure: may not apply if a founder species enters its very favourable adaptive zone. I 283 Species/Mayr: very conservative estimate of 10 million animal species, of which are ca. 1.5 million described. So about 15% known. Legitimate estimate: 30 million species. Only 5% are known. On the other hand, 99% of all bird species are discovered and described. In many insects, arachnids, low vertebrates probably less than 10%. The same applies to mushrooms, protists and prokaryotes. |
Mayr I Ernst Mayr This is Biology, Cambridge/MA 1997 German Edition: Das ist Biologie Heidelberg 1998 Gould I Stephen Jay Gould The Panda’s Thumb. More Reflections in Natural History, New York 1980 German Edition: Der Daumen des Panda Frankfurt 2009 Gould II Stephen Jay Gould Hen’s Teeth and Horse’s Toes. Further Reflections in Natural History, New York 1983 German Edition: Wie das Zebra zu seinen Streifen kommt Frankfurt 1991 Gould III Stephen Jay Gould Full House. The Spread of Excellence from Plato to Darwin, New York 1996 German Edition: Illusion Fortschritt Frankfurt 2004 Gould IV Stephen Jay Gould The Flamingo’s Smile. Reflections in Natural History, New York 1985 German Edition: Das Lächeln des Flamingos Basel 1989 |
Terminology | Gould | I 190 1. Stasis: most species show little change in one direction or another during their presence on Earth. 2. Sudden appearance: in all areas of life, species do not occur due to incessant changes in their predecessors, but suddenly and "fully developed". I 191 Evolution/Gould: evolution essentially proceeds in two ways: a) Definition phyletic transformation: an entire population changes from one state to another. If all evolutionary changes were to occur in this way, life would not last long. (See Evolution/Gould). b) Definition speciation: new species branch off from existing ones. All speciation theories assume that splits occur quickly in very small populations. Most theoreticians prefer the "allopatric" speciation (which happens in a different place). This is the orthodox view. With the "sympatric" speciation, new forms appear within the distribution area of the previous form. I 198 Definition preadaption: preadaption is derived from the thesis that other functions would be fulfilled in the initial stages, e.g. half a jaw could support the gills. Half a wing may have been used to catch prey, or to control body temperature. I 240 Definition Eozoon: an Eozoon is an early form of an animal. I 256 Definition protists: protists are single cell precursors. Definition Metazoen: a Metazoen is a multicellular offspring. I 258 Definition homologous similarity in common precursors: two organisms may have the same feature because they got it from a common ancestor. Definition analogous similarity: organisms with analogous similarity have no common precursors. The two organisms have a common feature that represents the result of a separate but similar evolutionary change in independent lines of development. I 281 Definition parallelism, Definition convergence: parallelism is a separate development of similar features in the course of evolution. This occurs very often. II 56 Definition diploid: animals with paired chromosomes in both sexes are diploid. Some animals use a different trait for sex determination: the females are diploid, but the males have instead of each female pair only one chromosome and are considered to be the first males. Definition haploid: organisms with only one chromosome (half of the diploid number) are haploid. In other words, the males develop ironically from unfertilized eggs and have no father. Fertilized eggs, on the other hand, produce diploid females. Animals using this system are called Definition Haplodiploid: the males develop from unfertilized eggs and have no father. Fertilized eggs, on the other hand, produce diploid females. This can be used to control the number of females. II 57 This fascinating system can help explain the origin of social systems in ants. Or also, for example, that a male mite dies before its own birth after fertilising its sisters in the womb. At least 10% of all known animal species are haplodiploid. II 186 Definition homeotic mutation: legs or parts of legs replace a variety of structures on the head mainly antennae and parts of the mouth. Not all incorrectly placed parts are homoeoses. William Bateson (not Gregory), who later invented the word genetics, called cases only homeotically in which organs that have the same development or evolutionary origin are replaced. II 192 Viable homoeostats that emulate the primordial forms are not really reborn ancestors. Double elements are formed, no old patterns are found. II 193 These things make it clear how few genes are responsible for regulating the basic order in the body of a fruit fly. II 240 Definition zoocentric: zoocentrism is a erspective that derives general principles from the behaviour of other animals and then completely subsumes the human being into this category, because we are undoubtedly also animals. Definition anthropocentric: a point of view is anthropocentric when it tries to subsume nature in us by considering our peculiarities as the goal of life from the very beginning. The zoocentric view can be extended to the caricature, which is often referred to as "nothing but error": the human is "nothing more than an animal" (reductionism). Popular science is flooding us with the excessively broad version of zoocentrism. II 331 Definition "genetic drift"/Gould: the genetic drift is the process of random increase or decrease of the gene frequency. II 352 Definition Clade: a clade is a branch on an evolutionary tree. Cladism tries to establish the branching pattern for a number of related species. II 353 Definition sister group: the sister group forms an upside-down Ypsilon: two tribes sharing a common ancestor from which no other tribe branches off. Gorillas and chimpanzees form a sister group. We can then consider the chimpanzee gorilla group as a unit and ask which primate forms the sister group with it. II 354 Definition derived feature: properties that only occur for members of a direct lineage are derived features. For example, all mammals have hair, which is not the case with any other vertebrate. II 355 Hair is a derived feature for the class of mammals, because it has developed only once in the common ancestors of mammals and therefore identifies a true branch in the family tree of vertebrates. Common derived characteristics are common to two or more strains and can be used to identify sister groups. II 356 GouldVsCladism: most derived features are ambiguous: they either tend to be too easily delimitable, or they are adaptive enough to be developed by several strains through natural selection independently of each other. II 360 Definition classification (cladism): classification was designed for the purpose of reflecting relative dimensions of similarity. Definition phenetism: phenetism is another theory of classification, it focuses solely on the overall similarity and tries to evade the reproach of subjectivity by referring to a large number of features, all of which are expressed numerically and processed by the computer. II 374 Definition "Telegony": Telegony means that features of long extinct ancestors reappear. They are also called "descendants from afar." Telegony refers to the idea that a producer could influence offspring that were not conceived by him. Definition "Pangenesis" (1868, provisionally developed by Darwin) thesis: all cells of the body produce small particles called "Gemmulae", which circulate throughout the body, accumulate in the gametes and eventually transfer the features to the offspring. GouldVsPangenesis: since the "Gemmulae" can change, acquired features can be inherited, which would be Lamarckism. II 377 Definition orthogenesis: orthogenesis is the assumption that a pre-drawn path is followed. IV 103 Doctrine of uniformity: (represented by Charles Lyell and James Hutton) the uppermost layers of the earth have remained unchanged for millions of years. IV 153 Definition Monogeny: (19th century): thesis: Monogeny is the assumption of a common ancestry of all humans from the ancestors Adam and Eve. (Lower races were later degenerated from original perfection.) Definition Polygeny: (19th century): thesis: Polygeny is the assumption that Adam and Eve are only the ancestors of the white peoples. IV 159 Definition subspecies: a subspecies is a population inhabiting a specific geographical area. IV 357 Definition sympatric: sympatric means being in the same place. Definition allopatric: allopatric means being in separate places (assuming that species can only develop separately). III 19 The "Full House": Gould's central argument: natural reality is an accumulation of individuals in populations. Variation is not reducible but "real" in the sense that "the world" consists of it. Error: it is wrong to always describe populations (according to Plato) as "average", which is then considered "typical". III 67 "Full House": "Full House" describes the need to focus not only on an abstract measure of an average or a central tendency, but on the variation within whole systems. Error: it is an error to consider the likely outcome for a single individual as a measure of a central tendency. |
Gould I Stephen Jay Gould The Panda’s Thumb. More Reflections in Natural History, New York 1980 German Edition: Der Daumen des Panda Frankfurt 2009 Gould II Stephen Jay Gould Hen’s Teeth and Horse’s Toes. Further Reflections in Natural History, New York 1983 German Edition: Wie das Zebra zu seinen Streifen kommt Frankfurt 1991 Gould III Stephen Jay Gould Full House. The Spread of Excellence from Plato to Darwin, New York 1996 German Edition: Illusion Fortschritt Frankfurt 2004 Gould IV Stephen Jay Gould The Flamingo’s Smile. Reflections in Natural History, New York 1985 German Edition: Das Lächeln des Flamingos Basel 1989 |